diff --git a/src/Generators/DurableTaskSourceGenerator.cs b/src/Generators/DurableTaskSourceGenerator.cs
index 25566bc2..89d868a2 100644
--- a/src/Generators/DurableTaskSourceGenerator.cs
+++ b/src/Generators/DurableTaskSourceGenerator.cs
@@ -1101,10 +1101,6 @@ static void AddActivityFunctionDeclaration(StringBuilder sourceBuilder, DurableT
             string inputType = activity.GetInputTypeForNamespace(targetNamespace);
             string outputType = activity.GetOutputTypeForNamespace(targetNamespace);
             string inputParameter = inputType + " input";
-            if (inputType.EndsWith("?", StringComparison.Ordinal))
-            {
-                inputParameter += " = default";
-            }
 
             string simplifiedActivityTypeName = SimplifyTypeName(activity.TypeName, targetNamespace);
             // GeneratedActivityContext is a generated class that we use for each generated activity trigger definition.
diff --git a/test/Generators.Tests/AzureFunctionsTests.cs b/test/Generators.Tests/AzureFunctionsTests.cs
index 3165af59..49d2f9bb 100644
--- a/test/Generators.Tests/AzureFunctionsTests.cs
+++ b/test/Generators.Tests/AzureFunctionsTests.cs
@@ -203,12 +203,11 @@ await TestHelpers.RunTestAsync<DurableTaskSourceGenerator>(
     [InlineData("int", "string")]
     [InlineData("string", "int")]
     [InlineData("Guid", "TimeSpan")]
+    [InlineData("object?", "string")]
     public async Task Activities_ClassBasedSyntax(string inputType, string outputType)
     {
-        // Nullable reference types need to be used for input expressions
-        string defaultInputType = TestHelpers.GetDefaultInputType(inputType);
-
         string code = $@"
+#nullable enable
 using System;
 using System.Threading.Tasks;
 using Microsoft.DurableTask;
@@ -219,11 +218,13 @@ public class MyActivity : TaskActivity<{inputType}, {outputType}>
     public override Task<{outputType}> RunAsync(TaskActivityContext context, {inputType} input) => Task.FromResult<{outputType}>(default!);
 }}";
 
-        // Build the expected InputParameter format (matches generator logic)
-        string expectedInputParameter = inputType + " input";
+        // The generated CallMyActivityAsync helper appends "= default" for nullable inputs because the
+        // input is followed by an optional options parameter. The [ActivityTrigger] function does not,
+        // because its input is followed by required parameters (see issue #730).
+        string expectedCallInputParameter = inputType + " input";
         if (inputType.EndsWith('?'))
         {
-            expectedInputParameter += " = default";
+            expectedCallInputParameter += " = default";
         }
 
         string expectedOutput = TestHelpers.WrapAndFormat(
@@ -233,13 +234,13 @@ public class MyActivity : TaskActivity<{inputType}, {outputType}>
 /// Calls the <see cref=""MyActivity""/> activity.
 /// </summary>
 /// <inheritdoc cref=""TaskOrchestrationContext.CallActivityAsync(TaskName, object?, TaskOptions?)""/>
-public static Task<{outputType}> CallMyActivityAsync(this TaskOrchestrationContext ctx, {inputType} input, TaskOptions? options = null)
+public static Task<{outputType}> CallMyActivityAsync(this TaskOrchestrationContext ctx, {expectedCallInputParameter}, TaskOptions? options = null)
 {{
     return ctx.CallActivityAsync<{outputType}>(""MyActivity"", input, options);
 }}
 
 [Function(nameof(MyActivity))]
-public static async Task<{outputType}> MyActivity([ActivityTrigger] {expectedInputParameter}, string instanceId, FunctionContext executionContext)
+public static async Task<{outputType}> MyActivity([ActivityTrigger] {inputType} input, string instanceId, FunctionContext executionContext)
 {{
     ITaskActivity activity = ActivatorUtilities.GetServiceOrCreateInstance<MyActivity>(executionContext.InstanceServices);
     TaskActivityContext context = new GeneratedActivityContext(""MyActivity"", instanceId);